Abstract
An updated revision of Oriental Dryinidae is presented. Seven subfamilies, 20 genera and 368 species are treated. Eight new species are described: Aphelopus zonalis Xu, Olmi & He, sp. nov. (China, Hainan); Anteon zoilum Xu, Olmi & He, sp. nov. (China, Yunnan), Anteon zonarium Xu, Olmi & He, sp. nov. (China, Yunnan), Anteon zopyrum Xu, Olmi & He, sp. nov. (China, Xizang), Anteon zoroastrum Xu, Olmi & He, sp. nov. (Malaysia, Malaya), Esagonatopus sinensis Xu, Olmi & He, sp. nov. (China, Yunnan), Gonatopus yunnanensis Xu, Olmi & He, sp. nov. (China, Yunnan); Ponomarenkoa ellenbergeri Olmi, Xu & He, sp. nov. (Myanmar amber). Descriptions, geographic distribution, known hosts, natural en-emies and type material of each species are presented, together with illustrations of the main morphological characters and keys to the subfamilies, genera and species. Complete lists of references concerning the Oriental Dryinidae and their hosts are given. New synonymies are proposed for Aphelopus albiclypeus Xu, He & Olmi, 1999 (=A. exnotaulices He & Xu, 2002, syn. nov.), A. orientalis Olmi, 1984 (=A. albopictoides Xu & He, 1999, syn. nov.), A. taiwanensis Olmi, 1991 (=A. compresssus Xu & Yao, 1997, syn. nov.), A. niger Xu & He, 1999 (=A. nigricornis Xu, He & Olmi, 1999, syn. nov.), A. penanganus Olmi, 1984 (=A.olmii He & Xu, 2002, syn. nov.), Anteon cacumen Xu & He, 1997 (=A. longwangshanense Xu & He, 1997, syn. nov.), A. hilare Olmi, 1984 (=A. corax Olmi, 1984, syn. nov., =A. javanum Olmi, 1984, syn. nov., =A. serratum Xu & He, 1999, syn. nov.), A. lankanum Olmi, 1984 (=A. planum Xu & He, 1999, syn. nov.), A. munitum Olmi, 1984 (=A. bauense Olmi, 1984, syn. nov.), A. parapriscum Olmi, 1991 (=A. alpinum He & Xu, 2002, syn. nov.), A. peterseni Olmi, 1984 (=A. scrupulosum He & Xu, 2002, syn. nov.), A. yuani Xu, He & Olmi, 1998 (=A. yuae He & Xu, 2002, syn. nov.), Lonchodryinus bimaculatus Xu & He, 1994 (=L. niger He & Xu, 2002, syn. nov.), L. ruficornis (Dalman, 1818) (=L. melaphelus Xu & He, 1994, syn. nov.), Dryinus indicus (Kieffer, 1914) (=Chlorodryinus koreanus Móczár, 1983, syn. nov., =Dryinus masneri Olmi, 2009, syn. nov.), D. stantoni Ashmead, 1904 (=D. undatomarginis Xu & He, 1998, syn. nov., =D. wuyishanensis He & Xu, 2002, syn. nov.), Adryinus jini Xu & Yang, 1995 (=A. platycornis Xu & He, 1995, syn. nov.), Gonatopus nigricans (R. Perkins, 1905 (=G. fulgori Nakagawa, 1906, syn. nov., =G. insulanus He & Xu, 1998, syn. nov., Pseudogonatopus sogatea Rohwer, 1920, syn. nov.; P. pusanus Olmi, 1984, syn. nov.), G. nudus (R. Perkins, 1912) (=G. yangi He & Xu, 1998, syn. nov.), G. pedestris Dalman, 1818 (=Epigonatopus sakaii Esaki & Hashimoto, 1933, syn. nov.), G. rufoniger Olmi, 1993 (=Neodryinus hishimonovorus Xu & He, 1997, syn. nov.), G. schen-klingi Strand, 1913 (=G. euscelidivorus Xu & He, 1999, syn. nov.). New combinations are proposed for Deinodryinus con-strictus (Olmi, 1998), comb. nov. (from Anteon), Dryinus asiaticus (Olmi, 1984), comb. nov. (from Alphadryinus), D. barbarus (Olmi, 1984), comb. nov. (from Mesodryinus), Gonatopus bengalensis (Olmi, 1984), comb. nov. (from Agona-topoides ), G. bicuspis (Olmi, 1993), comb. nov. (from Pseudogonatopus), G. borneanus (Olmi, 1984), comb. nov. (from Agonatopoides ); G. indicus (Olmi, 1987), comb. nov. (from Donisthorpina), G. insularis (Olmi, 1984), comb. nov. (from Agonatopoides), G. lankae (Ponomarenko, 1981), comb. nov. (from Pseudogonatopus), G. malesiae (Olmi, 1984), comb. nov. (from Pseudogonatopus), G. nepalensis (Olmi, 1986), comb. nov. (from Pseudogonatopus), G. pajanensis (Olmi, 1989), comb. nov. (from Agonatopoides), G. pyrillae (Mani, 1942), comb. nov. (from Agonatopoides), G. sarawakensis (Olmi, 1984), comb. nov. (from Pseudogonatopus), G. validus (Olmi, 1984), comb. nov. (from Pseudogonatopus).
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Olmi, M. (1995f) Contribution to the knowledge of the world Dryinidae (Hymenoptera Chrysidoidea). Phytophaga, 6, 3–54.
Olmi, M. (1995g) Olmi, M. (1995f) Hymenoptera Bethylidae, Dryinidae, Sclerogibbidae, Embolemidae. In: Minelli, A., Ruffo, S. & La Posta, S. (Eds.), Checklist delle specie della fauna italiana, 100. Calderini, Bologna, Italy, pp. 1–8.
Olmi, M. (1996a). Eucamptonyx garcetei n. sp., nuova specie di Dryinidae del Paraguay (Hymenoptera Chrysidoidea). Bollettino della Società entomologica italiana, Genova, 128, 65–68.
Olmi, M. (1996b) Systematics of the genera Echthrodelphax and Mercetia (Hymenoptera: Dryinidae). Memoirs of the Entomological Society of Washington, 17, 179–182.
Olmi, M. (1996c) New Anteoninae from Taiwan (Hymenoptera: Dryinidae). Oriental Insects, 30, 171–180. http://dx.doi.org/10.1080/00305316.1996.10433839
Olmi, M. (1996d) Taxonomic remarks on American Dryinidae, with descriptions of new species (Hymenoptera Chrysidoidea). Redia, 79, 57–81.
Olmi, M. (1998a) A contribution to the knowledge of the Dryinidae (Hymenoptera, Chrysidoidea). Lambillionea, XCVIII, 49–59.
Olmi, M. (1998b) Taxonomic remarks on Australian Dryinidae with description of new species (Insecta, Hymenoptera, Chrysidoidea). Spixiana, 21(1), 53–58.
Olmi, M. (1998c) A contribution to the knowledge of Dryinidae (Hymenoptera: Chrysidoidea) and Strepsiptera of Mozambique. Oriental Insects, 32, 59–78. http://dx.doi.org/10.1080/00305316.1998.10433767
Olmi, M. (1998d) Gonatopus orangensis: a new species of Dryinidae from South Africa (Hymenoptera Chrysidoidea). Tropical Zoology, 11, 177–181. http://dx.doi.org/10.1080/03946975.1998.10539359
Olmi, M. (1998e [“1997”]) New Embolemidae and Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica, N. S., 20 (33), 30–118.
Olmi, M. (1998f [“1997”]) New Oriental and Neotropic Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica, N. S., 20(33), 152–167.
Olmi, M. (1999a) Hymenoptera Dryinidae-Embolemidae. Fauna d'Italia, 37. Edizioni Calderini, Bologna, Italy, 425 pp.
Olmi, M. (1999b) I Driinidi e la loro utilizzazione in lotta biologica: problemi e prospettive. In: Gervasini, E. & Sala, A. (Ed.), Atti Convegno su “Metcalfa pruinosa: diffusione nel continente europeo e prospettive di controllo biologico”, S. Donato Milanese, 21 ottobre 1999, Sherwood-Foreste ed Alberi Oggi, 55 (Supplemento), 27–34.
Olmi, M. (2000a) Bio-ecologia degli Imenotteri Driinidi e loro impiego in programmi di lotta biologica. In: Lucchi, A. (Ed.), La Metcalfa negli ecosistemi italiani. ARSIA, Firenze, pp. 93–117.
Olmi, M. (2000b [“1998”]) New species of Oriental Anteon, with new synonymies (Hymenoptera Dryinidae). Frustula entomologica, N.S., 21 (34), 68–74.
Olmi, M. (2003 [“2001”]) A contribution to the knowledge of Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica, N.S., 24 (37), 21–49.
Olmi, M. (2004a) A contribution to the knowledge of Dryinidae (Hymenoptera: Chrysidoidea) of Northern Mozambique. Oriental Insects, 38, 353–372. http://dx.doi.org/10.1080/00305316.2004.10417403
Olmi, M. (2004b) Dryinidae e Sclerogibbidae raccolti nelle piccole isole dell'arcipelago toscano, sulla costa toscana, in Corsica e in Sardegna nel corso del progetto INTERREG II (Hymenoptera: Chrysidoidea). Resumés du IVe Congrés International sur Environnement et Identité en Méditerranée, Corte (Corsica), 19–25 Juillet 2004, 117.
Olmi, M. (2004c) Dryinidae (Hym.) from the Kuril Islands, Kamchatka and Sakhalin (Russia), with the description of a new species of Aphelopus Dalman. The Entomologist's Monthly Magazine, 140, 301–306.
Olmi, M. (2005a) A contribution to the knowledge of Oriental Dryinidae (Hymenoptera: Chrysidoidea), with descriptions of two new species from India and Thailand. Oriental Insects, 39, 345–352. http://dx.doi.org/10.1080/00305316.2005.10417447
Olmi, M. (2005b [“2004”]) Dryinidae e Sclerogibbidae raccolti nelle piccole isole dell'arcipelago toscano, sulla costa toscana, in Corsica e in Sardegna nel corso del progetto INTERREG II (Insecta: Hymenoptera: Chrysidoidea). Atti della Società Toscana di Scienze naturali, Memorie, Serie B, CXI, 121–127.
Olmi, M. (2006 [“2005”]) A catalogue of Dryinidae and Embolemidae of South Africa, with descriptions of new species (Hymenoptera Chrysidoidea). Frustula entomologica,N.S., 28–29(41–42), 1–57.
Olmi, M. (2007) New species of Afrotropical Dryinidae (Hymenoptera: Chrysidoidea), with description of a new genus and a new subfamily. African Invertebrates, 48(2), 199–232.
Olmi, M. (2008a) Order Hymenoptera, family Dryinidae. In: A. van Harten (Ed.), Arthropod fauna of the UAE, 1. Dar Al Ummah Printing, Abu Dhabi, pp. 361–371.
Olmi, M. (2008b) Descriptions of two new species of Dryinidae (Hym.) from Laos and India, with remarks on Anteon clavatum Olmi & Currado. Entomologist's Monthly Magazine, 144, 151–155.
Olmi, M. (2008c) Descriptions of new species of Dryinidae (Hymenoptera: Chrysidoidea) from Thailand. Oriental Insects, 42, 227–235. http://dx.doi.org/10.1080/00305316.2008.10417547
Olmi, M. (2009a) A contribution to the study of the Palaearctic Dryinidae, including descriptions of two new species from Japan and South Korea (Hymenoptera: Chrysidoidea). Entomologist's Gazette, 60 (2), 127–134.
Olmi, M. (2009b) New species of South African Dryinidae (Hymenoptera: Chrysidoidea). African Invertebrates, 50 (2), 447–460. http://dx.doi.org/10.5733/afin.050.0212
Olmi, M. (2010 [2008]) A contribution to the knowledge of Dryinidae from the Oriental, Nearctic, Neotropical and Australian regions (Hymenoptera Chrysidoidea). Frustula entomologica, N. S., 31 (44), 11–34.
Olmi, M. (2011 [2009]) A contribution to the knowledge of world Dryinidae (Hymenoptera Chrysidoidea). Frustula entomologica, N. S., 32(45), 43–76.
Olmi, M. (2012 [2010]) Description of Thaumatodryinus sharkeyi, new species of Dryinidae from Malaysia (Hymenoptera: Chrysidoidea). Frustula entomologica, N.S., 33 (46), 96–101.
Olmi, M. & Bechly, G. (2001) New parasitic wasps from Baltic amber (Insecta: Hymenoptera: Dryinidae). Stuttgarter Beiträge zur Naturkunde, Serie B, Nr. 306, 1–58.
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Olmi, M. & Currado, I. (1976) Gonatopodinae extraeuropei conservati nel Museo Civico di Storia Naturale di Genova (Hymenoptera Dryinidae). Annali del Museo Civico di Storia Naturale di Genova, 81, 165–194.
Olmi, M. & Currado, I. (1977) On the identity of Gonatopus pedestris Dalman (Hymenoptera: Dryinidae). Entomologica Scandinavica, 8, 76–78. http://dx.doi.org/10.1163/187631277X00125
Olmi, M. & Currado, I. (1979a) Anteoninae conservati nel Museo Civico di Storia Naturale di Genova (Hymenoptera Dryinidae). Annali del Museo Civico di Storia Naturale di Genova, 82, 340–349.
Olmi, M. & Currado, I. (1979b) Revisione del genere Haplogonatopus R.C.L. Perkins (Hymenoptera Dryinidae). Annali della Facoltà di Scienze Agrarie dell'Università di Torino, 11, 37–44.
Olmi, M. & De Rond, J. (2001) Dryinidae. In: Dathe, H., Taeger, A. & Blank, S.M. (Eds.), Verzeichnis der Hautflügler Deutschlands. Entomofauna Germanica, 4, pp. 115–116.
Olmi, M. & Guglielmino, A. (2010a) Description of Erwiniinae, new subfamily of Dryinidae from Ecuador (Hymenoptera: Chrysidoidea). Zootaxa, 2605, 56–62.
Olmi, M. & Guglielmino, A. (2010b) Revision of Nearctic species of Lonchodryinus Kieffer 1905, with description of a new species from New Mexico (Hymenoptera: Dryinidae). Zootaxa, 2654, 30–40.
Olmi, M. & Guglielmino, A. (2010c) Revision of Nearctic species of Esagonatopus, with description of a new species from Florida (Hymenoptera, Dryinidae). ZooKeys, 70, 57–66. http://dx.doi.org/10.3897/zookeys.70.764
Olmi, M. & Guglielmino, A. (2011) Revision of fossil species of Dryinus belonging to lamellatus group, with description of a new species (Hymenoptera, Dryinidae). ZooKeys, 130, 505–514. http://dx.doi.org/10.3897/zookeys.130.1335
Olmi, M., Guglielmino, A. & Vollaro, M. (2011) Revision of fossil species of Dryinus belonging to constans group, with description of a new species (Hymenoptera: Dryinidae). Zootaxa, 2981, 43–55.
Olmi, M., Rasnitsyn, A.P. & Guglielmino, A. (2010) Revision of rock fossils of Dryinidae and Embolemidae (Hymenoptera: Chrysidoidea). Zootaxa, 2499, 21–38.
Olmi, M., Virla, E.G. & Fernández, F., 2000. Las Avispas Dryinidae de la Región Neotropical (Hymenoptera: Chrysidoidea). Biota Colombiana, 1 (2), 141–163.
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